![]() 1A) and he was also senior author with JC Meakins and JG Priestley of the 1918/1919 paper. The author of the 1915 paper was John Scott Haldane (Fig. The importance of the ratio, VA/QC, in pulmonary gas exchange did not appear in this paper de novo but emerged in two papers, published by physiologists in 19, concerned with the volume of the respiratory dead space. The concept was first suggested in a paper published in the 1918/1919 volume of The Journal of Physiology to account for the crippling breathlessness suffered by the soldiers who had survived gas attacks in the First World War. Large differences in the ratio of alveolar ventilation to alveolar capillary blood flow, VA/QC, in different regions of the lungs have been recognised as a principal cause of hypoxaemia in respiratory disease since the 1950s.
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